Jeffrey Buler

Headshot of Jeffrey Buler
Department of Entomology and Wildlife Ecology

Jeffrey Buler

Associate Professor of Wildlife Ecology

Office location:
531 South College Avenue
146 Townsend Hall
Newark, DE 19716


  • Ph.D. Biology, The University of Southern Mississippi, 2006
  • M.S. Wildlife, Louisiana State University , 1999
  • B.A. Biology , St. Mary’s College of Maryland, 1995

Regular courses taught

  • ENWC 416/616, Wildlife Habitat Management, Fall
  • ENWC325, Wildlife Management, Alternate Springs
  • ENWC 820 Landscape Ecology, Alternate Springs
  • ENWC 842 Ecological Modeling, Alternate Springs


  • Radar Aeroecology
  • Avian Ecology and Migration
  • Landscape Ecology
  • Conservation Biology

12. Calderon, Lily. In progress. A multiscale approach to understanding the function and management of stopover habitats for migratory land birds

11. Cabrera-Cruz, Sergio. In progress. Nocturnal light pollution and landbird migration. Ph.D. Entomology and Wildlife Ecology

10. Clipp, Hannah. 2018. An investigation of landbird stopover patterns along the U.S. coast of the Gulf of Mexico during spring migration. M.S. Wildlife Ecology

9. Zenzal, T.J. 2018. Application of integrated long term datasets to understand migrant-habitat use along the Mississippi-Alabama coast. Post-doctoral Researcher.

8. Schreckengost, Timothy. 2017. Methods for studying stopover ecology of migrating landbirds with weather surveillance radar. M.S. Wildlife Ecology. Current position: General manager of Phoneskope.

7. McLaren, James. 2015-2016. Assess possible impacts of Superstorm Sandy on migrating landbirds and their habitats. Post-doctoral Researcher. Current position: Post-doctoral Researcher Environment Canada, Ontario, Canada.

6. Boone, Matthew. 2016. The effect of Hurricane Sandy on landbird migration in the Northeast United States. M.S. Wildlife Ecology. Current position: GIS/database manager Kauai Endangered Seabird Recovery Project, HI.

5. Adalsteinsson, Solny. 2016. (co-advised with W. G. Shriver). The effect of forest fragment dynamics on human disease risk: developing a holistic approach to prevent tick-borne diseases in urban landscapes. Ph.D. Entomology and Wildlife Ecology. 2016 UD CANR William J. Benton Graduate Student Award. Current position: post-doctoral researcher Washington University, MO.

4. Horton, Kyle. 2013. Examining songbird migration along the Delaware Bay: A comprehensive approach. M.S. in Wildlife Ecology. Current position: Ph.D student, University of Oklahoma, OK.

3. Sieges, Mason. 2013. If You Flood It, They Will Come: Quantifying Waterbird Response to the Migratory Bird Habitat Initiative. M.S. Wildlife Ecology. Current position: Biologist, Edwin B. Forsythe National Wildlife Refuge, NJ.

2. La Puma, David. 2013. Radar analysis of bird migration stopover sites in the southeastern U.S. Post-docoral Researcher. Current position: Director of Cape May Bird Observatory, NJ.

1. Duren, Ken. 2010. A predictive habitat model for northern bobwhite (Colinus virginianus) in Delaware. M.S. Wildlife Ecology. (co-advised with Dr. Chris Williams). Current position: Biometrician, Pennsylvania Department of Conservation and Natural Resources, PA.

Generally, JJB  will appear as second or last author if publication is a mentored student project. Google Scholar reported citations: 644, h-Index = 12, i10-Index = 14. Google Scholars Report Page


28. Ladin, Z. S., S. Van Nieuland, S. A. Adalsteinsson, V. D’Amico, J. L. Bowman, J. J. Buler, J. M. Baetens, B. De Baets, W. G. Shriver. 2018. Post-fledging space use of Nearctic-Neotropical migratory birds within an urbanized landscape. Movement Ecology. 

27. Adalsteinsson, S. A., J. J. Buler, J. L. Bowman, V. D’Amico, Z. Ladin, W. G. Shriver. 2018. Post‐independence mortality of juveniles is driven by anthropogenic hazards for two passerines in an urban landscape. Journal of Avian Biology.

26. Cabrera-Cruz, S. A., J. A. Smolinsky, J. J. Buler. 2018. Light pollution is brightest during the migratory phase of the annual cycle for nocturnally migrating birds around the world. Scientific Reports 8:3261.

25. Adalsteinsson, S. A., W. G. Shriver, A. Hojgaard, J. L. Bowman, D. Brisson, and J. J. Buler. 2018. Multiflora rose invasion amplifies prevalence of Lyme disease pathogen, but not necessarily Lyme disease risk. Parasites & Vectors 11:54.

24. McLaren, J. D., J. J. Buler (co-first author), T. Schreckengost, J. A. Smolinsky, M. Boone, E. E. van Loon, D. K. Dawson, E. L. Walters. 2018. Artificial light confounds broad-scale habitat use by migrating birds. Ecology Letters 21:356-364.

23. Zenzal, T. J., Smith, R. J., D. N. Ewert, R. H. Diehl, and J. J. Buler. 2018. Fine-scale heterogeneity drives forest use by spring migrant landbirds across a broad, contiguous forest matrix. The Condor: Ornithological Applications 120:166-184.


22. Buler, J. J., W. Barrow Jr., M. Boone, D. K. Dawson, R. H. Diehl, F. Moore, J. Smolinksky, T. Schreckengost. 2017. Linking Animals Aloft with the Terrestrial Landscape. pp 347-378 in: Chilson P., Frick W., Kelly J., Liechti F. (eds) Aeroecology. Springer, Cham.

21. Buler J. J., R. J. Lyon*, J. A. Smolinsky, T. J. Zenzal, and F.R. Moore. 2017. Body mass and wing shape explain variability in broad-scale bird species distributions along an ecological barrier during migratory stopover. Oecologia 185:205-212.

20. La Sorte, F., D. Fink, J. J. Buler, A. Farnsworth, and S. Cabrera-Cruz. 2017. Seasonal associations with urban light pollution for nocturnally migrating bird populations. Global Change Biology.

19. Cohen, E. B., W. C. Barrow, Jr., J. J. Buler, J. L. Deppe, A. Farnsworth, P. P. Marra, S. R. McWilliams, D. W. Mehlman, R. R. Wilson, M. S. Woodrey, and F. R. Moore. 2017. Understanding how en route events around the Gulf of Mexico impact intercontinental migratory landbird populations. The Condor: Ornithological Applications 119(2):327-343.

18. Archibald, K. M., J. J. Buler, J. A. Smolinsky, and R. J. Smith. 2017. Migrating birds reorient toward land at dawn over the Great Lakes, USA. The Auk 134:193201.


17. Adalsteinsson, S. A, V. D’Amico, W. G. Shriver, D. Brisson, and J. J. Buler. 2016. Scale-dependent effects of nonnative plant invasion on host-seeking tick abundance. Ecosphere 7.

16. Lafleur, J. M., and J. J. Buler (co-first author), and F. R. Moore. 2016. Geographic position and landscape composition explain regional patterns of migrating landbird distributions during spring stopover along the northern coast of the Gulf of Mexico. Landscape Ecology 31:1697–1709.

15. Horton, K. G., W. G. Shriver, and J. J. Buler. 2016. An assessment of spatio-temporal relationships between nocturnal bird migration traffic rates and diurnal bird stopover density. Movement Ecology 4:1–10.


14. Horton, K. G., W. G. Shriver, and J. Buler. 2015. A Comparison of traffic estimates of nocturnal flying animals using radar, thermal imaging, and acoustic recording.  Ecological Applications 25:390-401.


13. Buler, J. J., and D. K. Dawson. 2014.  Radar analysis of fall bird migration stopover sites in the northeastern US. The Condor 116:357–370.

12. Sieges, M. L., J. J. Buler, J. A. Smolinsky, W. Barrow Jr., M. Baldwin, and L. Randall. 2014. Assessment of bird response to Migratory Bird Habitat Initiative managed wetlands using weather surveillance radar. Southeastern Naturalist 13: G36–G65.


11. Irvin, E., K. Duren, J. Buler. W. Jones, A, Gonzon, and C. K. Williams. 2013. A multi-scale occupancy model for the grasshopper sparrow in the Mid-Atlantic. Journal of Wildlife Management.


10. Buler, J.J., L.A. Randall, J.P. Fleskes, W.C. Barrow, T. Bogart, and D. Kluver. 2012. Mapping wintering waterfowl distributions using weather surveillance radar. PLoS ONE 7:e41571

9. Duren, K. R., J. J. Buler, W. Jones, and C. K. Williams. 2012. Effects of broadcasting calls during surveys to estimate density and occupancy of northern bobwhite. Wildlife Society Bulletin 36:16-20.


8. Duren, K., J. J. Buler, W. Jones, and C. K. Williams. 2011. An improved multi-scale approach to modeling habitat occupancy of northern bobwhite. Journal of Wildlife Management 75:1700–1709.

7. Buler, J. J., and F. R. Moore. 2011. Migrant – habitat relations during stopover along an ecological barrier: Extrinsic constraints and conservation implications. Journal of Ornithology 152(S1):101–112.


6. Buler, J. J. and R. H. Diehl. 2009. Quantifying bird density during migratory stopover using weather surveillance radar. IEEE Transcations on Geoscience and Remote Sensing 47:2741–2751.


5. Ruth, J. M., J. J. Buler, R. H. Diehl, and R. S. Sojda, R.S. 2008. Management and research applications of long-range surveillance radar data for birds, bats, and flying insects. U.S. Geological Survey Fact Sheet 2008-3095, 4p.


4. Barrow, W., J. J. Buler, B. Couvillion, R. Diehl, S. Faulkner, F. Moore, L. Randall. 2007. Broad scale response of landbird migration to the immediate effects of hurricane Katrina. Science and the storms – the USGS response to the hurricanes of 2005. (G.S. Farris, G. J. Smith, M. P Crane, C. R. Demas, L. L. Robbins, D. L. Lavoire, eds.). U.S. Geological Survey Circular 1306. pp. 131-136.

3. Buler, J. J., F. R. Moore, and S. Woltmann. 2007. A multi-scale examination of stopover habitat use by birds. Ecology 88:1789–1802.


2. Moore, F. R., M. S. Woodrey, J. J. Buler, S. Woltmann, and T. R. Simons. 2005. Understanding the stopover of migratory birds: A scale dependent approach. Bird Conservation Implementation and Integration in the Americas: Proceedings of the Third International Partners in Flight Conference 2002 March 20-24; Asilomar, California, Vol 2. (C.J. Ralph and T.D. Rich, eds.). Pacific Southwest Research Station, Forest Service, U.S.D.A., Gen. Tech. Rep. PSW-191, Albany, CA pp. 684–689.


1. Buler, J. J., and R. B. Hamilton. 2000. Predation of natural and artificial nests in a southern pine forest. The Auk 117:739–747.