Charles Bartlett

Department of Entomology and Wildlife Ecology

Charles Bartlett

Associate Professor of Entomology

Office location:
531 South College Avenue
250 Townsend Hall
Newark, DE 19716


  • Ph.D., Systematic Entomology, North Carolina State University, 1996.
  • M.S., Entomology and Applied Ecology, University of Delaware, 1992.
  • B.S., Wildlife Management, Forestry minor, University of Vermont, 1987.

Current courses

  • Insect Taxonomy (ENWC 406/606)
  • Field Taxonomy (ENWC 408/608)
  • Insect Structure and Function (ENWC 405/605)
  • Larval Taxonomy (ENWC 602)

Research activities

  • Primary interest in the alpha-taxonomy of delphacid planthoppers (Hemiptera: Fulgoroidea: Delphacidae), particularly of the New World.  More broadly interested in the taxonomy, biodiversity, host relations, and biogeography of planthoppers.
  • Director of University of Delaware Insect Reference Collection.  The University of Delaware collection contains ca. 350,000 pinned specimens plus ~500 slides and ~2,500 alcohol specimens.  I am responsible for all aspects of the collection including outgoing and incoming loans, accessions, visitors, curation, and general organization.

Professional activities

  • American Entomological Society (Executive Council, Recording Secretary since 1998).
  • Entomological Society of America.
  • Xerces Society.
  • Organization of Tropical Studies.
  • Entomological Collections Network
  • Society for the Preservation of Natural History Collections

6. Hagerty, Tyler. In progress. Investigation of Anastatus (Eupelmidae, Hymenoptera) as potential biocontrol agents of Lycorma delicatula (Fulgoridae) and other Hemiptera including potential non-target organisms.

5. Gonzon, Anthony T. In progress. A Revision of the Polyphyletic Genus Delphacodes Fieber, 1866 (Hemiptera: Delphacidae) in the New World.

4. Kennedy, Ashley C. 2013. A Revision of the Genus Caenodelphax Fennah (Hemiptera: Fulgoroidea: Delphacidae).

3. Weglarz, Kathryn. 2012. A Revision of the Genus Chionomus Fennah (Hemiptera: Fulgoroidea: Delphacidae): Cleaning out Delphacodes Fieber.

2. Barringer, Lawrence. 2012. Canopy assemblages and species richness of plant hoppers (Fulgoroidea) in the Amazon Canopy of Ecuador.

1. Donovall, Rick. 2007. Generic Revision and Phylogenetic Analysis of the Dictyopharid Planthopper Subfamily Dictyopharinae (Hemiptera: Fulgoroidea) in the New World. M.S. in Entomology.


57. Bartlett, C. R. 2019. A new genus and species of Delphacidae (Hemiptera: Fulgoroidea: Delphacidae) from Costa Rica. Zootaxa 4657 (2): 361–368.

56. Huang, Yi-Xin, Feng-Juan Ren, C. R. Bartlett, Yong-Sheng Wei and Dao-Zheng Qin. 2019 Contribution to the mitogenome diversity in Delphacinae: Phylogenetic and ecological implications. Genomics (in Press, ePub ahead of paper)

55. Wang, Wenqian, Yixin Huang, C. R. Bartlett, Fanmei Zhou, Rui Meng and Daozheng Qin. 2019 Characterization of the complete mitochondrial genomes of two species of the genus Aphaena Guérin-Méneville (Hemiptera: Fulgoridae) and its phylogenetic implications. International Journal of Biological Macromolecules 141: 29-40.

54. Constant, J. and C. R. Bartlett. 2019. New records and species in five planthopper families from Keo Seima Wildlife Sanctuary, Cambodia with checklist of Cambodian planthoppers (Hemiptera: Fulgoromorpha). Belgian Journal of Entomology 83: 1–27.

53. Otero, M., and C. R. Bartlett. 2019. A new species of Abbrosoga (Hemiptera: Fulgoroidea: Delphacidae), an endemic Puerto Rican planthopper genus, with an updated checklist of the Delphacidae of Puerto Rico. Zootaxa 4563(2): 372-386.

52. Wallner, A. M. and C. R. Bartlett. 2019. Comparative morphology of female gonapophyses IX in Delphacidae (Hemiptera: Auchenorrhyncha: Fulgoromorpha) with key to tribes. Zootaxa 4564 (1): 137–172.

51. Bahder, B.W., C.R. Bartlett, E. A. Barrantes Barrantes, M.A. Zumbado Echavarria, A. R. Humphries, E.E. Helmick, M.S. Ascunce, E.M. Goss. 2019. A new species of Omolicna (Hemiptera: Auchenorrhyncha: Fulgoroidea: Derbidae) from coconut palm in Costa Rica and new country records for Omolicna brunnea and Omolicna triata. Zootaxa 4577 (3): 501–514.


50. Gonçalves da Silva, F., E. M. dos Passos, L. E. C. Diniz, M. F. Fernandes, C. R. Bartlett, M. Dollet and A. V. Teodoro. 2018. Relative contribution of rainfall and coconut hybrids to the abundance and composition of the Auchenorrhyncha community as potential vectors of phytoplasmas in the state of Sergipe, Brazil. Agricultural and Forest Entomology 21: 190-198 (ePub, early view 27 Dec. 2018), 9 pp. 10.1111/afe.12320

49. Hill, J.G., J.L. Seltzer, A. Hendon and C.R. Bartlett. 2018. First Report of the American Palm Cixiid (Hemiptera: Cixiidae) from Mississippi, USA. Transactions of the American Entomological Society 144(3): 593-597.

48. Barringer, L. E. and C. R. Bartlett. 2018. Pennsylvania planthoppers (Hemiptera: Auchenorrhyncha: Fulgoroidea): relative abundance and incidental catch using novel trapping methods. Insecta Mundi 0661: 1–31.

47. Bartlett, C. R., E. M. dos Passos. F. G. da Silva, L. E. C. Diniz and M. Dollet. 2018. A new species of Oecleus Stål (Hemiptera: Fulgoroidea: Cixiidae) from coconut in Brazil. Zootaxa 4472 (2): 358–364.

46. Bartlett, C. R. and A. C. Kennedy. 2018. A review of New World Malaxa (Hemiptera: Fulgoroidea: Delphacidae). Zootaxa 4441(3): 511–528.

45. Bartlett, C. R., L. L. Deitz, D. A. Dmitriev, A. F. Sanborn, A. Soulier-Perkins, M. S. Wallace. 2018. The Diversity of the True Hoppers (Hemiptera: Auchenorrhyncha). Pp. 501-590. In: R. G. Foottit and P. H. Adler. Insect Biodiversity II. Wiley-Blackwell, Chichester, United Kingdom. Xxxv + 987 + 21 plates.

44. Zhi-Shun Song, C. R. Bartlett, L. B. O’Brien and Ai-Ping Liang.  2018. Morphological phylogeny of the family Dictyopharidae (Hemiptera: Fulgoromorpha), with emphasis on the relationships among the tribes of Dictyopharinae. Systematic Entomology

43. Gnezdilov, V. M. and C. R. Bartlett. 2018. A new genus and two new species of the family Issidae (Hemiptera, Auchenorrhyncha: Fulgoroidea) from Amazonian Ecuador.  Proceedings of the Entomological Society of Washington 120:62–75.


42. Leavengood, J. M., C. R. Bartlett, and S. Vitanza-Hedman. 2107. First reports of six planthoppers (Hemiptera: Fulgoroidea: Tagosodes, Delphacodes, Pareuidella, Nilaparvata, Asarcopus, Bruchomorpha) in Texas. Entomological News 127:215-229.

41. Huang, Yi-Xin; Li-Fang Zheng; C. R. Bartlett; Dao-Zheng Qin. 2017. Resolving phylogenetic relationships of Tropidocephalini (Hemiptera: Delphacidae: Delphacinae) as inferred from four genetic loci. Scientific Reports 7 (3319): 10 pp. doi:10.1038/s41598-017-03624-w. (

40. Seltmann, K. C., N. S. Cobb, L. F. Gall, C. R. Bartlett, M. A. Basham, I. Betancourt, C. Bills, B. Brandt, R. L. Brown, C. Bundy, M. S. Caterino, C. Chapman, A. Cognato, J. Colby, S. P. Cook, K. M. Daly, L. Dyer, N. M. Franz, J. K. Gelhaus, C. C. Grinter, C. E. Harp, R. L. Hawkins, S. L. Heydon, G. M. Hill, S. Huber, N. Johnson, A. Y. Kawahara, L. S. Kimsey, B. C. Kondratieff, F.-T. Krell, L. Leblanc, S. Lee, C. J. Marshall, L. M. McCabe, J. V. McHugh, K. L. Menard, P. A. Opler, N. Palffy-Muhoray, N. Pardikes, M. A. Peterson, N. E. Pierce, A. Poremski, D. S. Sikes, J. D. Weintraub, D. Wikle, J. M. Zaspel, G. Zolnerowich.  2017. LepNet: The Lepidoptera of North America Network. ZooTaxa 4247 (1): 073-077.


39. Wheeler, A. G., Jr., E. R. Hoebeke and C. R. Bartlett. 2016. Juncus effusus (Juncaceae) as a host plant of Nothodelphax occlusa (Van Duzee) (Hemiptera: Fulgoroidea: Delphacidae) in the Pacific Northwest. Proceedings of the Entomological Society of Washington 18(4):629-635.

38. Gnezdilov, V.M., C. R. Bartlett and T. Bourgoin. 2016. A new tribe of Tropiduchidae (Hemiptera, Fulgoroidea) with revision of the genus Buca Walker and description of asymmetric hind leg spinulation. Florida Entomologist 99(3): 406-416.

37. Llano, C.A., C. R. Bartlett and G. Guevarra. 2016. First record of the subfamily Asiracinae and Copicerus irroratus (Hemiptera: Auchenorrhyncha: Delphacidae) in Colombia. Florida Entomologist 99(1):120-122.


36. Bartlett, C. R. 2015. No Kelisia Fieber (Hemiptera: Fulgoroidea: Delphacidae) in South America: New taxonomic placement of Phrictopyga vittata (Muir) comb. nov. from Brazil. Entomological News 125(4): 289-292.

35. Kennedy, A. C. and C. R. Bartlett. 2015. Scientific Note: An unusual morphotype of Caenodelphax teapae (Fowler) (Hemiptera: Fulgoroidea: Delphacidae). Entomological News 125(3): 224-227.

34. Bartlett, C. R., S. W. Wilson and D. S. Sikes. 2015. First New World record of Paradelphacodes paludosus (Flor 1861) (Hemiptera: Fulgoroidea: Delphacidae) in Alaska. Entomological News 124(5): 370-371.

33. Wheeler, A. G., jr. and C. R. Bartlett. 2015. Megamelanus bicolor Ball (Hemiptera: Fulgoroidea: Delphacidae): a specialist planthopper on saltgrass (Distichlis spicata; Poaceae) in Nebraska’s saline wetlands. Proceedings of the Entomological Society of Washington 117(1): 45-54.

32. Bartlett, C. R., and G. Kunz. 2015. A new genus and species of delphacid planthopper (Hemiptera: Fulgoroidea: Delphacidae) from Central America with a preliminary regional species list. Zootaxa 3946:510–518.


31. Bartlett, C. R., and M. D. Webb. 2014. The planthopper genus Spartidelphax, a new segregate of Nearctic Delphacodes (Hemiptera, Delphacidae). ZooKeys 453: 19-36 (10 Nov 2014). doi: 10.3897/zookeys.453.8369.

30. Bartlett, C. R., L. B. O’Brien and S. W. Wilson.  2014.  A review of the planthoppers (Hemiptera: Fulgoroidea) of the United States. Memoirs of the American Entomological Society 50:1–287.

29. Bartlett, C. R. 2014. New species of the planthopper genus Parkana (Hemiptera: Fulgoroidea: Delphacidae) from Mesoamerica. Transactions of the American Entomological Society 140:185–208.

28. Kennedy, A. C. and C. R. Bartlett. 2014. Systematics of Caenodelphax Fennah (Hemiptera: Fulgoroidea: Delphacidae) and description of the new genus Flavoclypeus. Transactions of the American Entomological Society 140:17–65.


27. Kennedy, A. C., C. R. Bartlett and S. W. Wilson. 2012. An annotated checklist of the delphacid planthoppers (Hemiptera: Delphacidae) of Florida with the description of three new species and the new genus, Meristopsis. Florida Entomologist 95:395–421.


26. Barringer, L. E. and C. R. Bartlett. 2011. A review of New World Asiracinae (Hemiptera: Auchenorrhyncha: Delphacidae) with five new taxa. Cicadina 12:7–39.

25. Weglarz, K. M. and C. R. Bartlett. 2011. Akemetopon, a new genus containing three new species of planthoppers (Hemiptera: Fulgoroidea: Delphacidae). Zootaxa 3007:50–60.

24. Bartlett, C. R. and K.G.A. Hamilton. 2011. Aethodelphax prairianus gen. et sp. nov. (Hemiptera: Delphacidae) and seven congeneric species from North American DelphacodesZootaxa 2837:48–66.

23. Bartlett C. R., E. R. Adams, A. T. Gonzon. 2011. Planthoppers of Delaware (Hemiptera, Fulgoroidea), excluding Delphacidae, with species Incidence from adjacent States. ZooKeys 2011:1–42.


22. Urban, J. M., C. R. Bartlett, and J. R. Cryan. 2010. Evolution of Delphacidae (Hemiptera: Fulgoroidea): Combined-evidence phylogenetics reveals importance of grass host shifts. Systematic Entomology. 35: 678–691.


21. Bartlett, C. R. 2009. Diversity in New World Stenocranine Planthoppers (Hemiptera: Delphacidae). Transactions of the American Entomological Society 135:443–486.

20. Bartlett, C. R. 2009. A new genus of New World Tropidocephalini (Hemiptera: Delphacidae: Delphacinae) with two new species. Entomological News 120:443–452.

19. Wallace, M. S., C. R. Bartlett, L. L. Deitz, and M. J. Rothschild. 2009. New State Records of Treehoppers (Hemiptera: Membracidae) for Delaware, Maryland, Mississippi, Pennsylvania, and West Virginia. Proceedings of the Entomological Society of Washington. 111:734–737.


18. Bartlett, C. R. 2008. Case 3460. Liburnia Stål, 1866 (Insecta, Hemiptera, Delphacidae): proposed designation of Embolophora monoceros Stål, 1855 as the type species. Bulletin of Zoological Nomenclature. 65:183–187.

17. Gonzon, A. T., and C. R. Bartlett. 2008. Systematics of Hadropygos n.g., Metadelphax Wagner and New World Toya Distant (Hemiptera: Delphacidae). Transactions of the American Entomological Society 113:205–277.

16. Bartlett, C. R., L. L. Deitz, M. J. Rothschild, and M. S. Wallace.  2008.  Treehopper diversity (Hemiptera: Membracidae) of Little Orleans, Allegany Co., Maryland. Proceedings of the Entomological Society of Washington 110:130–143.


15. Bartlett, C. R. 2007. A review of the planthopper genus Nilaparvata (Hemiptera: Delphacidae) in the New World.  Entomological News 118:49–66.

14. Bartlett, C. R., and A. G. Wheeler, Jr.  2007.  Kelisia and Stenocranus species (Hemiptera: Fulgoromorpha: Delphacidae): new host-plant associations and distributional records of eight sedge-feeding planthoppers.  Proceedings of the Entomological Society of Washington 109:400–415.


13. Gonzon, A. T., C. R. Bartlett, and J. L. Bowman. 2006. Planthopper (Hemiptera: Fulgoroidea) Diversity in the Great Smoky Mountains National Park. Transactions of the American Entomological Society 132:243–260.

12. Wheeler, A. G., jr., and C. R. Bartlett.  2006.  Delphacodes campestris (Van Duzee) and D. lutulenta (Van Duzee) (Hemiptera: Fulgoromorpha: Delphacidae): association with common hairgrass, Deschampsia flexuosa (Poaceae), and notes on habitats, seasonality, and taxonomy.  Proceedings of the Entomological Society of Washington 118:395–403.


11. Bartlett, C. R. 2005.  Two new genera and species of stenocranine planthoppers (Hemiptera: Delphacidae) from North America.  Entomological News 116:291–303.


10. Bartlett, C. R., and J. L. Bowman. 2004. Preliminary Inventory of the Planthoppers (Hemiptera:: Fulgoroidea) of the Great Smoky Mountains National Park, North Carolina and Tennessee, U.S.A. Entomological News 114:246–254.

9. Heckscher, C. M., and C. R. Bartlett. 2004. Rediscovery and habitat associations of Photuris bethaniensis (Coleoptera: Lampyridae). Coleopterists Bulletin 58:349–353.

8. Deitz, L. L., and C. R. Bartlett. 2004. Publications of Thomas Kenneth Wood (1942–2002). Proceedings of the Entomological Society of Washington 106: 586–591.


7. Bartlett, C. R. 2002. A new genus and species of delphacid planthoppers (Hemiptera: Fulgoroidea) from Canada. Entomological News 113:97–102.

6. Cryan, J. R., and C. R. Bartlett. 2002. A redescription of the Caribbean genus Antillotolania Ramos (Hemiptera: Membracidae), with two new species. Annals of the Entomological Society of America 95: 302–308.


5. Chordas Ill, S. W., C. R. Bartlett, and R. L. Stewart Jr. 2001. First backswimmer records (Hemiptera: Notonectidae) from Delaware. Entomological News 112:183–186.


4. Bartlett, C. R., and L. L. Deitz. 2000. Revisions of the New World Delphacid planthopper genus Pissonotus (Hemiptera: Fuloroidea). Thomas Say Publication in Entomology. Entomological Society of America, Lanham, Maryland.

3. Bartlett, C. R. 2000. An annotated list of planthoppers (Hemiptera:Fulgoroidea) of Guana Island (British West Indes). Entomological News 111:120–132.


2. McCann, J. M., S. E. Mabey, L. J. Niles, C. R. Bartlett, P. Kerlinger. 1993. A regional study of coastal migratory stopover habitat for Neotropical migrant songbirds: land management implications. Transactions of the North American Wildlife and Natural Resources Conference 58:398–407.


1. Johnson, R. K., R. R. Roth, S. E. Kleiner, and C. R. Bartlett. 1991. A case of polygyny in the wood thrush. The Wilson Bulletin 103:509–510